Queen Beryl is the first major villain in every version of the Sailor Moon series. She is the commander of the Dark Kingdom as well as the commander of the four Shitenou. During Silver Millennium, she led the invasion forces to conquer the Moon Kingdom.
She is generally portrayed in a more tragic light in the manga than she is in the anime. In the English-dubbed anime no mention is given to her origin and she is only said to be the ruler of the Negaverse, an evil alternate dimension, from which she was somehow able to escape in order to attack this dimension. At the end of the season, Beryl is said to be merely banished back to the Negaverse when Sailor Moon defeats her.
